Dr Alex Clarke

University position

Senior Research Associate

Dr Alex Clarke is pleased to consider applications from prospective PhD students.


Department of Psychology

Home page

https://sites.google.com/site/al... (personal home page)

Research Theme

Cognitive and Behavioural Neuroscience


How does the brain make sense of the world? We can recognise tens of thousands of objects, but despite this vast number, our recognition is remarkably quick and accurate, completed within a few hundred milliseconds. My research ask what are the neural dynamics and mechanisms by which vision activates semantics, and how does our knowledge about the world around us change these dynamics?

Through combining MEG, EEG, fMRI, Neuropsychology and Augmented Reality, we can take a multi-modal approach to gain a deeper understanding of the nature of semantic representations in the brain.

Increased recurrent interactions during specific object naming
Increased recurrent interactions during specific object naming
Click image to view full-size

Research Focus


Object Recognition

Conceptual Knowledge




Clinical conditions

No direct clinical relevance


Augmented Reality (AR)

Behavioural analysis

Electrocorticography (ECoG)

Electroencephalography (EEG)

Magnetic resonance imaging (MRI)

Magnetoencephalography (MEG)

Transcranial magnetic stimulation (TMS)



Alexandra Krugliak

Victoria Nicholls

Lorraine Tyler

Associated News Items



Clarke, A (2017), “Top-down cognitive influences on object recognition: a commentary on Perry and Lupyan (2016)” Language, Cognition and Neuroscience


Clarke A, Pell PJ, Ranganath C, Tyler LK (2016), “Learning warps object representations in the ventral temporal cortex” Journal of Cognitive Neuroscience 28(7): 1010-1023


Clarke A (2015), “Dynamic information processing states revealed through neurocognitive models of object semantics” Language, Cognition and Neuroscience 30:4: 409-419

Clarke A, Tyler LK (2015), “Understanding what we see: how we derive meaning from vision” TICS 19(11): 677-687

Wright P, Randall B, Clarke A, Tyler LK (2015), “The perirhinal cortex and conceptual processing: Effects of feature-based statistics following damage to the anterior temporal lobes.” Neuropsychologia Details


Clarke A, Devereux BJ, Randall B, Tyler LK (2014), “Predicting the Time Course of Individual Objects with MEG.” Cereb Cortex Details

Clarke A, Tyler LK (2014), “Object-specific semantic coding in human perirhinal cortex.” J Neurosci 34(14):4766-75 Details


Clarke A, Taylor KI, Devereux B, Randall B, Tyler LK (2013), “From perception to conception: how meaningful objects are processed over time.” Cereb Cortex 23(1):187-97 Details

Devereux BJ, Clarke A, Marouchos A, Tyler LK (2013), “Representational similarity analysis reveals commonalities and differences in the semantic processing of words and objects.” J Neurosci 33(48):18906-16 Details

Tyler LK, Cheung TP, Devereux BJ, Clarke A (2013), “Syntactic computations in the language network: characterizing dynamic network properties using representational similarity analysis.” Front Psychol 4:271 Details

Tyler LK, Chiu S, Zhuang J, Randall B, Devereux BJ, Wright P, Clarke A, Taylor KI (2013), “Objects and categories: feature statistics and object processing in the ventral stream.” J Cogn Neurosci 25(10):1723-35 Details


Clarke A, Taylor KI, Tyler LK (2011), “The evolution of meaning: spatio-temporal dynamics of visual object recognition.” J Cogn Neurosci 23(8):1887-99 Details


Andrews TJ, Clarke A, Pell P, Hartley T (2010), “Selectivity for low-level features of objects in the human ventral stream.” Neuroimage 49(1):703-11 Details